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The California State University

Publications

  • Hays, G., L. Ferreira, A. Sequeira, M. Meekan, C. Duarte, H. Bailey, F. Bailleul, W. Bowen, M. Caley and D. Costa. 2016. Key Questions in Marine Megafauna Movement Ecology. Trends in Ecology & Evolution 31: 463-475.

  • Kelsey, E., R. Bradley, P. Warzybok, J. Jahncke and S. Shaffer. 2016. Environmental temperatures, artificial nests, and incubation of Cassin's auklet. The Journal of Wildlife Management 80: 292-299.

  • Thompson, D., L. Torres, G. Taylor, M. Rayner, P. Sagar, S. Shaffer, R. Phillips and S. Bury. 2015. Stable isotope values delineate the non-breeding distributions of sooty shearwaters Puffinus griseus in the North Pacific Ocean. Marine Ecology Progress Series 521: 277-282.

  • Thorne, L., E. Hazen, S. Bograd, D. Foley, M. Conners, M. Kappes, H. Kim, D. Costa, Y. Tremblay and S. Shaffer. 2015. Foraging behavior links climate variability and reproduction in North Pacific albatrosses. Movement Ecology 3: 27.

  • Frechette, D., A. Osterback, S. Hayes, J. Moore, S. Shaffer, M. Pavelka, C. Winchell and J. Harvey. 2015. Assessing the Relationship between Gulls Larus spp. and Pacific Salmon in Central California Using Radiotelemetry. North American Journal of Fisheries Management 35: 775-788.

  • Kappes, M., S. Shaffer, Y. Tremblay, D. Foley, D. Palacios, S. Bograd and D. Costa. 2015. Reproductive constraints influence habitat accessibility, segregation, and preference of sympatric albatross species. Movement Ecology 3: 34.

  • Osterback, A., D. Frechette, S. Hayes, S. Shaffer and J. Moore. 2015. Long-term shifts in anthropogenic subsidies to gulls and implications for an imperiled fish. Biological Conservation 191: 606-613.

  • Orben, R., D. Irons, R. Paredes, D. Roby, R. Phillips, S. Shaffer and L. Manne. 2015. North or south? Niche separation of endemic red‐legged kittiwakes and sympatric black‐legged kittiwakes during their non‐breeding migrations. Journal of Biogeography 42: 401-412.

  • Gutowsky, S., M. Leonard, M. Conners, S. Shaffer and I. Jonsen. 2015. Individual-level Variation and Higher-level Interpretations of Space Use in Wide-ranging Species. Frontiers in Marine Science 2.

  • Orben, R., R. Paredes, D. Roby, D. Irons, S. Shaffer and J. Gill. 2015. Body size affects individual winter foraging strategies of thick‐billed murres in the Bering Sea. Journal of Animal Ecology 84: 1589-1599.

  • Orben, R., R. Paredes, D. Roby, D. Irons and S. Shaffer. 2015. Wintering North Pacific black-legged kittiwakes balance spatial flexibility and consistency. Movement Ecology 3: 36.

  • Conners, M., E. Hazen, D. Costa and S. Shaffer. 2015. Shadowed by scale. Movement Ecology 3: 28.

  • Young, H., S. Maxwell, M. Conners and S. Shaffer. 2015. Pelagic marine protected areas protect foraging habitat for multiple breeding seabirds in the central Pacific. Biological Conservation 181: 226-235.

  • Gutowsky, S., L. Gutowsky, I. Jonsen, M. Leonard, M. Naughton, M. Romano and S. Shaffer. 2014. Daily activity budgets reveal a quasi-flightless stage during non-breeding in Hawaiian albatrosses. Movement Ecology 2: 23.

  • Shaffer, S., C. Clatterbuck, E. Kelsey, A. Naiman, L. Young, E. VanderWerf, P. Warzybok, R. Bradley, J. Jahncke and G. Bower. 2014. As the Egg Turns. PLOS ONE 9: e97898-e97897.

  • Gutowsky, S., Y. Tremblay, M. Kappes, E. Flint, J. Klavitter, L. Laniawe, D. Costa, M. Naughton, M. Romano and S. Shaffer. 2014. Divergent post‐breeding distribution and habitat associations of fledgling and adult Black‐footed Albatrosses Phoebastria nigripes in the North Pacific. Ibis 156: 60-72.

  • Carey, M., R. Phillips, J. Silk and S. Shaffer. 2014. Trans-equatorial migration of Short-tailed Shearwaters revealed by geolocators. EMU 114: 352-359.

  • Osterback, A., D. Frechette, S. Hayes, M. Bond, S. Shaffer and J. Moore. 2014. Linking individual size and wild and hatchery ancestry to survival and predation risk of threatened steelhead (Oncorhynchus mykiss). Canadian Journal of Fisheries and Aquatic Sciences 71: 1877-1887.

  • Osterback, A., D. Frechette, A. Shelton, S. Hayes, M. Bond, S. Shaffer and J. Moore. 2013. High predation on small populations. Ecosphere 4: art116-art121.

  • Maxwell, S., E. Hazen, S. Bograd, B. Halpern, G. Breed, B. Nickel, N. Teutschel, L. Crowder, S. Benson and P. Dutton. 2013. Cumulative human impacts on marine predators. Nature Communications 4: 2688-2696.

  • Hazen, E., S. Jorgensen, R. Rykaczewski, S. Bograd, D. Foley, I. Jonsen, S. Shaffer, J. Dunne, D. Costa and L. Crowder. 2013. Predicted habitat shifts of Pacific top predators in a changing climate. Nature Climate Change 3: 234-238.

  • Frechette, D., A. Osterback, S. Hayes, M. Bond, J. Moore, S. Shaffer and J. Harvey. 2012. Assessing Avian Predation on Juvenile Salmonids using Passive Integrated Transponder Tag Recoveries and Mark-Recapture Methods. North American Journal of Fisheries Management 32: 1237-1250.

  • Rayner, M., G. Taylor, H. Gummer, R. Phillips, P. Sagar, S. Shaffer and D. Thompson. 2012. breeding cycle, year-round distribution and activity patterns of the endangered Chatham Petrel (Pterodroma axillaris). EMU 112: 107-116.

  • Winship, A., S. Jorgensen, S. Shaffer, I. Jonsen, P. Robinson, D. Costa and B. Block. 2012. State‐space framework for estimating measurement error from double‐tagging telemetry experiments. Methods in Ecology and Evolution 3: 291-302.

  • Verrier, D., C. Guinet, M. Authier, Y. Tremblay, S. Shaffer, D. Costa, R. Groscolas and J. Arnould. 2011. ontogeny of diving abilities in subantarctic fur seal pups. Functional Ecology 25: 818-828.

  • Rayner, M., G. Taylor, D. Thompson, L. Torres, P. Sagar and S. Shaffer. 2011. Migration and diving activity in three non‐breeding flesh‐footed shearwaters Puffinus carneipes. Journal of Avian Biology 42: 266-270.

  • Žydelis, R., R. Lewison, S. Shaffer, J. Moore, A. Boustany, J. Roberts, M. Sims, D. Dunn, B. Best and Y. Tremblay. 2011. Dynamic habitat models. Proceedings: Biological Sciences 278: 3191-3200.

  • Shaffer, S. 2011. review of seabird energetics using the doubly labeled water method. Comparative Biochemistry and Physiology, Part a 158: 315-322.

  • Rayner, M., M. Hauber, T. Steeves, H. Lawrence, D. Thompson, P. Sagar, S. Bury, T. Landers, R. Phillips and L. Ranjard. 2011. Contemporary and historical separation of transequatorial migration between genetically distinct seabird populations. Nature Communications.

  • Block et al., B.A. 2011. Tracking Apex Marine Predator Movements in a Dynamic Ocean. Nature 475: 86-90.

  • Shaffer, S.A. 2011. A review of seabird energetics using the doubly labeled water method. Comparative Biochemistry and Physiology 158: 315-322.

  • Raymond, B., S. Shaffer, S. Sokolov, E. Woehler, D. Costa, L. Einoder, M. Hindell, G. Hosie, M. Pinkerton and P. Sagar. 2010. Shearwater Foraging in the Southern Ocean. PLOS ONE 5: e10960-e10960.

  • Young, H., D. McCauley, R. Dirzo, R. Dunbar and S. Shaffer. 2010. Niche partitioning among and within sympatric tropical seabirds revealed by stable isotope analysis. Marine Ecology Progress Series 416: 285-294.

  • Young, H., S. Shaffer, D. McCauley, D. Foley, R. Dirzo and B. Block. 2010. Resource partitioning by species but not sex in sympatric boobies in the central Pacific Ocean. Marine Ecology Progress Series 403: 291-301.

  • Kappes, M., S. Shaffer, Y. Tremblay, D. Foley, D. Palacios, P. Robinson, S. Bograd and D. Costa. 2010. Hawaiian albatrosses track interannual variability of marine habitats in the North Pacific. Progress in Oceanography 86: 246-260.

  • Gonzalez-Solis, J. and S. Shaffer. 2009. Introduction and synthesis. Marine Ecology Progress Series 391: 117-120.

  • Shaffer, S., H. Weimerskirch, D. Scott, D. Pinaud, D. Thompson, P. Sagar, H. Moller, G. Taylor, D. Foley and Y. Tremblay. 2009. Spatiotemporal habitat use by breeding sooty shearwaters Puffinus griseus. Marine Ecology Progress Series 391: 209-220.

  • Adams, J., D. Scott, S. McKechnie, G. Blackwell, S. Shaffer and H. Moller. 2009. Effects of geolocation archival tags on reproduction and adult body mass of sooty shearwaters (Puffinus griseus). New Zealand Journal of Zoology 36: 355-366.

  • Tremblay, Y., S. Bertrand, R. Henry, M. Kappes, D. Costa and S. Shaffer. 2009. Analytical approaches to investigating seabird-environment interactions. Marine Ecology Progress Series 391: 153-163.

  • Weimerskirch, H., S. Shaffer, Y. Tremblay, D. Costa, H. Gadenne, A. Kato, Y. Ropert-Coudert, K. Sato and D. Aurioles. 2009. Species- and sex-specific differences in foraging behaviour and foraging zones in blue-footed and brown boobies in the Gulf of California. Marine Ecology Progress Series 391: 267-278.

  • Shaffer, S.A., H. Weimerskirch, D. Scott, D. Pinaud, D.R. Thompson, P.M. Sagar, H. Moller, G.A. Taylor, D.G. Foley, Y. Tremblay and D.P. Costa. 2009. Spatio-temporal Habitat Use by Breeding Sooty Shearwaters (Puffinus griseus). Marine Ecology Progress Series 391: 209-220.

  • Young, L., C. Vanderlip, D. Duffy, V. Afanasyev and S. Shaffer. 2009. Bringing Home the Trash. PLoS ONE.

  • Burger, A. and S. Shaffer. 2008. Perspectives in Ornithology Application of Tracking and Data-Logging Technology in Research and Conservation of Seabirds. The Auk 125: 253-264.

  • Suryan, R., D. Anderson, S. Shaffer, D. Roby, Y. Tremblay, D. Costa, P. Sievert, (...) and N. Nakamura. 2008. Wind, Waves, and Wing Loading. PLoS ONE.

  • Angelier, F., S. Shaffer, H. Weimerskirch, C. Trouvé and O. Chastel. 2007. Corticosterone and foraging behavior in a pelagic seabird. Physiological and Biochemical Zoology : PBZ.

  • Shaffer, S. and D. Costa. 2006. database for the study of marine mammal behavior. IEEE Journal of Oceanic Engineering 31: 82-86.

  • Tremblay, Y., S. Shaffer, S. Fowler, C. Kuhn, B. McDonald, M. Weise, C. Bost, H. Weimerskirch, D. Crocker and M. Goebel. 2006. Interpolation of animal tracking data in a fluid environment. Journal of Experimental Biology 209: 128-140.

  • Shaffer, S., G. Gabrielsen, J. Verreault and D. Costa. 2006. Validation of Water Flux and Body Composition in Glaucous Gulls (Larus hyperboreus). Physiological & Biochemical Zoology 79: 836-845.

  • Shaffer, S., Y. Tremblay, H. Weimerskirch, D. Scott, D. Thompson, P. Sagar, H. Moller, G. Taylor, D. Foley, B. Block and D. Costa. 2006. Migratory shearwaters integrate oceanic resources across the Pacific Ocean in an endless summer. Proceedings of the National Academy of Sciences of the United States of America 103: 12799-12802.

  • Tremblay, Y., S. Shaffer, S. Fowler, C. Kuhn, B. McDonald, M. J.Weise, C. Bost, H. Weimerskirch, D. Crocker, M. Goebel and D. Costa. 2006. Interpolation of animal tracking data in a fluid environment. Journal of Experimental Biology 209: 128-140.

  • Angelier, F., S. Shaffer, H. Weimerskirch and O. Chastel. 2006. Effect of age, breeding experience and senescence on corticosterone and prolactin levels in a long-lived seabird. General & Comparative Endocrinology 149: 1-9.

  • Kuhn, C., B. McDonald, S. Shaffer, J. Barnes, D. Crocker, J. Burns and D. Costa. 2006. Diving physiology and winter foraging behavior of a juvenile leopard seal ( Hydrurga leptonyx). Polar Biology 29: 303-307.

  • Shaffer, S., Y. Tremblay, J. Awkerman, R. Henry, S. Teo, D. Anderson, D. Croll, B. Block and D. Costa. 2005. Comparison of light- and SST-based geolocation with satellite telemetry in free-ranging albatrosses. Marine Biology 147: 833-843.

  • Mabille, G., O. Boutard, S. Shaffer, D. Costa and H. Weimerskirch. 2004. Growth and energy expenditure of Wandering Albatross Diomedea exulans chicks. Ibis 146: 85-94.

  • Shaffer, S., D. Costa and H. Weimerskirch. 2004. Field Metabolic Rates of Black-Browed Albatrosses Thalassarche melanophrys during the Incubation Stage. Journal of Avian Biology 35: 551-558.

  • Shaffer, S. 2004. Annual energy budget and food requirements of breeding wandering albatrosses (Diomedea exulans). Polar Biology 27: 253-256.

  • Costa, D., C. Kuhn, M. Weise, S. Shaffer and J. Arnould. 2004. When does physiology limit the foraging behaviour of freely diving mammals? International Congress Series 1275: 359-366.

  • Costa, Daniel P, Scott A Shaffer and CALIFORNIA UNIV SANTA CRUZ. 2003. Database for the Study of Marine Mammal Behavior. Ft. Belvoir

  • Weimerskirch, H., S. Shaffer, G. Mabille, J. Martin, O. Boutard and J. Rouanet. 2002. Heart rate and energy expenditure of incubating wandering albatrosses. Journal of Experimental Biology 205: 475-483.

  • Shaffer, S., D. Costa and H. Weimerskirch. 2001. Comparison of Methods for Evaluating Energy Expenditure of Incubating Wandering Albatrosses. Physiological and Biochemical Zoology: Ecological and Evolutionary Approaches 74: 823-831.

  • Shaffer, S., D. Costa and H. Weimerskirch. 2001. Behavioural Factors Affecting Foraging Effort of Breeding Wandering Albatrosses. Journal of Animal Ecology 70: 864-874.

  • Shaffer, S.A. 2000. Foraging Ecology of Wandering Albatrosses (Diomedea exulans): Impacts on reproduction and life history. PhD diss., University of California Santa Cruz.

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Educational Background

  • 1996-2000 – Ph.D., University of California Santa Cruz (Biology)

  • 1994-1996 – M.Sc., University of California Santa Cruz (Marine Science)

  • 1988-1993 – B.Sc., San Diego State University (Biology)

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Collaborators, Advisors & Advisees

  • Block, B – Hopkins Marine Station, Stanford University

  • Conners, M – University of California Santa Cruz

  • Costa, D – University of California Santa Cruz

  • Gabrielsen, G – Norwegian Polar Institute

  • Hyrenbach, D – Hawaii Pacific University

  • Orben, R – University of California Santa Cruz

  • Suryan, R – Oregon Statte University

  • Weimerskirch, H – CEBC - CNRS, France

  • Young, H – Stanford University

  • Young, L – Pacific Rim Conservation

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