Skip to side navigation Skip to content
The California State University

Publications

  • Ikeda, C., W. Cochlan, C. Bronicheski, V. Trainer, C. Trick and J. Raven. 2016. effects of salinity on the cellular permeability and cytotoxicity of Heterosigma akashiwo. Journal of Phycology 52: 745-760.

  • Mansfeldt, C., L. Richter, B. Ahner, W. Cochlan, R. Richardson and S. Chen. 2016. Use of De Novo transcriptome libraries to characterize a novel oleaginous marine Chlorella species during the accumulation of triacylglycerols. PLoS ONE.

  • Greene, C., M. Huntley, I. Archibald, L. Gerber, D. Sills, J. Granados, J. Tester, C. Beal, M. Walsh and R. Bidigare. 2016. Marine Microalgae. Oceanography 29: 10-15.

  • McCabe, R., B. Hickey, R. Kudela, K. Lefebvre, N. Adams, B. Bill, F. Gulland, R. Thomson, W. Cochlan and V. Trainer. 2016. unprecedented coastwide toxic algal bloom linked to anomalous ocean conditions. Geophysical Research Letters 43: 10-10,376.

  • Wells, M., V. Trainer, T. Smayda, B. Karlson, C. Trick, R. Kudela, A. Ishikawa, S. Bernard, A. Wulff' and D. Anderson. 2015. Harmful algal blooms and climate change. Harmful Algae 49: 68-93.

  • Bartal, R., B. Shi, W. Cochlan and E. Carpenter. 2015. model system elucidating calcification functions in the prymnesiophyteEmiliania huxleyireveals dependence of nitrate acquisition on coccoliths. Limnology and Oceanography 60: 149-158.

  • Auro, M. and W. Cochlan. 2013. Nitrogen Utilization and Toxin Production by Two Diatoms of thePseudo-nitzschia pseudodelicatissimaComplex. Journal of Phycology 49: 156-169.

  • Bittar, T., Y. Lin, L. Sassano, B. Wheeler, S. Brown, W. Cochlan and Z. Johnson. 2013. Carbon allocation under light and nitrogen resource gradients in two model marine phytoplankton. Journal of Phycology 49: 523-535.

  • Trainer, V., S. Bates, N. Lundholm, A. Thessen, W. Cochlan, N. Adams and C. Trick. 2012. Pseudo-nitzschia physiological ecology, phylogeny, toxicity, monitoring and impacts on ecosystem health. Harmful Algae: 271-300.

  • Trick, C., B. Bill, W. Cochlan, M. Wells, V. Trainer and L. Pickell. 2010. Iron enrichment stimulates toxic diatom production in high-nitrate, low-chlorophyll areas. Proceedings of the National Academy of Sciences 107: 5887-5892.

  • Kudela, R., S. Seeyave and W. Cochlan. 2010. role of nutrients in regulation and promotion of harmful algal blooms in upwelling systems. Progress in Oceanography: 1-2.

  • Trick, C., B. Bill, W. Cochlan, M. WeIIs, V. Trainer and L. Pickell. 2010. Iron enrichment stimulates toxic diatom production in high-nitrate, low-chlorophyll areas. Proceedings of the National Academy of Sciences of the United States of America 107: 5887-5892.

  • Pickell, L., M. Wells, C. Trick and W. Cochlan. 2009. sea‐going continuous culture system for investigating phytoplankton community response to macro‐ and micro‐nutrient manipulations. Limnology and Oceanography: Methods 7: 21-32.

  • Trainer, V., B. Hickey, E. Lessard, W. Cochlan, C. Trick, M. Wells, A. MacFadyen and S. Moore. 2009. Variability of Pseudo‐nitzschia and domoic acid in the Juan de Fuca eddy region and its adjacent shelves. Limnology and Oceanography 54: 289-308.

  • Pickell, L., M. Wells, C. Trick and W. Cochlan. 2009. sea-going continuous culture system for investigating phytoplankton community response to macro- and micro-nutrient manipulations. Limnology and Oceanography: Methods 7: 21-32.

  • Trainer, V., M. Wells, W. Cochlan, C. Trick, B. Bill, K. Baugh, B. Beall, J. Herndon and N. Lundholmf. 2009. ecological study of a massive bloom of toxigenic Pseudo‐nitzschia cuspidata off the Washington State coast. Limnology and Oceanography 54: 1461-1474.

  • Wells, M., M. Perry, W. Cochlan and C. Trick. 2009. Effects of Iron Complexing Ligands on the Long Term Ecosystem Response to Iron Enrichment of HNLC waters. .

  • Wells, M., C. Trick, W. Cochlan and B. Beall. 2009. Persistence of iron limitation in the western subarctic Pacific SEEDS II mesoscale fertilization experiment. Deep Sea Research Part II: Topical Studies in Oceanography.

  • Kudo, I., Y. Noiri, W. Cochlan, K. Suzuki, T. Aramaki, T. Ono and Y. Nojiri. 2009. Primary productivity, bacterial productivity and nitrogen uptake in response to iron enrichment during the SEEDS II. Deep Sea Research Part II: Topical Studies in Oceanography.

  • Trainer, V., M. Wells, W. Cochlan, C. Trick, B. Bill, K. Baugh, B. Beall, J. Herndon and N. Lundholm. 2009. ecological study of a massive bloom of toxigenic Pseudo-nitzschia cuspidata off the Washington State coast. Limnology & Oceanography 54: 6-6.

  • Cochlan, W. 2008. Nitrogen Uptake in the Southern Ocean-Chapter 12. Nitrogen in the Marine Environment: 569-596.

  • Olson, M., E. Lessard, W. Cochlan and V. Trainer. 2008. Intrinsic Growth and Microzooplankton Grazing on Toxigenic Pseudo-nitzschia spp. Diatoms from the Coastal Northeast Pacific. Limnology and Oceanography 53: 1352-1368.

  • Anderson, D., J. Burkholder, W. Cochlan, P. Glibert, C. Gobler, C. Heil, R. Kudela, M. Parsons, J. Rensel and D. Townsend. 2008. Harmful algal blooms and eutrophication. Harmful Algae 8: 39-53.

  • Glibert, P., R. Azanza, M. Burford, K. Furuya, E. Abal, A. Al-Azri, F. Al-Yamani, (...) and M. Zhu. 2008. Ocean urea fertilization for carbon credits poses high ecological risks. Marine Pollution Bulletin.

  • Kudela, R., J. Lane and W. Cochlan. 2008. potential role of anthropogenically derived nitrogen in the growth of harmful algae in California, USA. Harmful Algae.

  • Cochlan, W. 2008. Nitrogen Uptake in the Southern Ocean. Nitrogen in the Marine Environment, 2nd Ed. D.G. Capone, D.A. Bronk, M.R. Mulholland, & E.J. Carpenter [Eds.]: 569-596.

  • Cochlan, W., J. Herndon and R. Kudela. 2008. Inorganic and organic nitrogen uptake by the toxigenic diatom Pseudo-nitzschia australis (Bacillariophyceae). Harmful Algae.

  • Anderson, D., J. Burkholder, W. Cochlan, P. Glibert, C. Gobler, C. Heil, R. Kudela, (...) and G. Vargo. 2008. Harmful algal blooms and eutrophication. Harmful Algae.

  • Tsuda, A., S. Takeda, H. Saito, J. Nishioka, I. Kudo, Y. Nojiri, K. Suzuki, M. Uematsu, M. Wells and D. Tsumune. 2007. Evidence for the grazing hypothesis. Journal of Oceanography 63: 983-994.

  • Howard, M., W. Cochlan, N. Ladizinsky and R. Kudela. 2007. Nitrogenous preference of toxigenic Pseudo-nitzschia australis (Bacillariophyceae) from field and laboratory experiments. Harmful Algae.

  • Trainer, V., W. Cochlan, A. Erickson, B. Bill, F. Cox, J. Borchert and K. Lefebvre. 2007. Recent domoic acid closures of shellfish harvest areas in Washington State inland waterways. Harmful Algae.

  • Herndon, J. and W. Cochlan. 2007. Nitrogen utilization by the raphidophyte Heterosigma akashiwo. Harmful Algae.

  • Kudela, R., W. Cochlan, T. Peterson and C. Trick. 2006. Impacts on phytoplankton biomass and productivity in the Pacific Northwest during the warm ocean conditions of 2005. Geophysical Research Letters 33: L22S06-L22S05.

  • Wells, M., C. Trick, W. Cochlan, M. Hughes and V. Trainer. 2005. Domoic acid. Limnology & Oceanography 50: 1908-1917.

  • Coale, K., K. Johnson, F. Chavez, K. Buesseler, R. Barber, M. Brzezinski, W. Cochlan, F. Millero, P. Falkowski and J. Bauer. 2004. Southern Ocean Iron Enrichment Experiment. Science 304: 408-414.

  • Cochlan, W., D. Bronk and K. Coale. 2002. Trace metals and nitrogenous nutrition of Antarctic phytoplankton. Deep Sea Research Part II: Topical Studies in Oceanography.

  • Cochlan, W. 2001. Heterotrophic Bacterial Response during a Mesoscale Iron Enrichment Experiment (IronEx II) in the Eastern Equatorial Pacific Ocean. Limnology and Oceanography 46: 428-435.

  • Cochlan, W. and D. Bronk. 2001. Nitrogen uptake kinetics in the Ross Sea, Antarctica. Deep Sea Research Part II: Topical Studies in Oceanography.

  • Kudela, R. and W. Cochlan. 2000. Nitrogen and carbon uptake kinetics and the influence of irradiance for a red tide bloom off southern California. Aquatic Microbial Ecology 21: 31-47.

  • Kudela, R., W. Cochlan and R. Dugdale. 1997. Carbon and nitrogen uptake response to light by phytoplankton during an upwelling event. Journal of Plankton Research 19: 609-630.

  • Berges, J., W. Cochlan and P. Harrison. 1995. Laboratory and field responses of algal nitrate reductase to diel periodicity in irradiance, nitrate exhaustion, and the presence of ammonium. Marine Ecology Progress Series 124: 259-269.

  • Cochlan, W., J. Wikner, G. Steward, D. Smith and F. Azam. 1993. Spatial distribution of viruses, bacteria and chlorophyll a in neritic, oceanic and estuarine environments. Marine Ecology Progress Series 92: 77-87.

  • Suttle, C., J. Stockner and W. Cochlan. 1991. Size-Dependent Ammonium and Phosphate Uptake, and N. Canadian Journal of Fisheries and Aquatic Sciences 48: 1226-1234.

  • Cochlan, W. and P. Harrison. 1991. Kinetics of nitrogen (nitrate, ammonium and urea) uptake by the picoflagellate Micromonas pusilla (Prasinophyceae). Journal of Experimental Marine Biology and Ecology.

  • Cochlan, W. and P. Harrison. 1991. UPTAKE OF NITRATE, AMMONIUM, AND UREA BY NITROGEN-STARVED CULTURES OF MICROMONAS PUSILLA (PRASINOPHYCEAE). Journal of Phycology 27: 673-679.

  • Cochlan, W. and P. Harrison. 1991. Inhibition of nitrate uptake by ammonium and urea in the eucaryotic picoflagellate Micromonas pusilla (Butcher) Manton et Parke. Journal of Experimental Marine Biology and Ecology.

  • Stockner, J., M. Klut and W. Cochlan. 1990. Leaky Filters. Canadian Journal of Fisheries and Aquatic Sciences 47: 16-23.

  • Cochlan, W. 1986. Seasonal study of uptake and regeneration of nitrogen on the Scotian Shelf. Continental Shelf Research.

return to top

Educational Background

  • 1990 – Ph.D., University of British Columbia, Vancouver, BC (Oceanography)

  • 1982 – M.Sc., Dalhousie University, Halifax, NS (Oceanography)

  • 1978 – B.Sc. (H), University of British Columbia, Vancouver, BC (Marine Biology)

return to top