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The California State University


  • Bean, W.T., D. Tange and S.D. Osborn. 2016. A suitability model for white-footed voles with insight into habitat associations at the southern boundary of their range. Northwestern Naturalist 97: 105-112.

  • Dulava, S., W. Bean and O. Richmond. 2015. Applications of unmanned aircraft systems (UAS) for waterbird surveys. Environmental Practice 17: 201.

  • Bean, W., L. Prugh, R. Stafford, H. Butterfield, M. Westphal, J. Brashares and P. Stephens. 2014. Species distribution models of an endangered rodent offer conflicting measures of habitat quality at multiple scales. Journal of Applied Ecology 51: 1116-1125.

  • Bean, W., R. Stafford, H. Butterfield and J. Brashares. 2014. Multi-Scale Distribution Model for Non-Equilibrium Populations Suggests Resource Limitation in an Endangered Rodent. PLOS ONE 9: e106638-e106638.

  • Bean, W. 2012. Spatial Ecology of the Giant Kangaroo Rat (Dipodomys ingens). .

  • Bean, W., R. Stafford, L. Prugh, H. Butterfield and J. Brashares. 2012. Evaluation of Monitoring Methods for the Endangered Giant Kangaroo Rat. Wildlife Society Bulletin 36: 587-593.

  • Bean, W., R. Stafford and J. Brashares. 2012. Effects of small sample size and sample bias on threshold selection and accuracy assessment of species distribution models. Ecography 35: 250-258.

  • Prugh, L., C. Stoner, C. Epps, W. Bean, W. Ripple, A. Laliberte and J. Brashares. 2009. Rise of the Mesopredator. BioScience 59: 779-791.

  • Bean, W. and E. Sanderson. 2008. Using a spatially explicit ecological model to test scenarios of fire use by Native Americans. Ecological Modelling 211: 301-308.

  • Wittemyer, G., P. Elsen, W. Bean, A. Burton and J. Brashares. 2008. Accelerated Human Population Growth at Protected Area Edges. Science 321: 123-126.

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Educational Background

  • 2007-2012 – Ph.D., UC-Berkeley (Environmental Science, Policy & Management)

  • 1999-2004 – B.A., Columbia University (Ecology, Evolution and Environmental Biology)

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